Quarterly report pursuant to Section 13 or 15(d)

Related-Party Transactions - Additional Information (Detail)

v3.20.1
Related-Party Transactions - Additional Information (Detail) - USD ($)
3 Months Ended
Mar. 31, 2020
Mar. 31, 2019
Cellectis [Member]    
Related Party Transaction [Line Items]    
Minimum net worth required $ 300,000,000  
Cellectis [Member] | License and Royalty Fees [Member]    
Related Party Transaction [Line Items]    
Related party transaction, expenses from transactions with related party 33,000 $ 25,000
University of Minnesota [Member] | License, Milestone Payments, Royalties and Legal Fees [Member]    
Related Party Transaction [Line Items]    
Related party transaction, expenses from transactions with related party 12,000 4,000
General and Administrative Expense [Member] | Cellectis [Member]    
Related Party Transaction [Line Items]    
Related party transaction, expenses from transactions with related party $ 62,000 $ 361,000